Dandelion pollen grains, macro shot of a bumblebee head and poppy flower

Selected publications

For a complete and most up-to-date list please visit Google Scholar or Researchgate. For PDFs of papers please email

Leah Salm (2023) Community gardens in Brighton- report of online survey results. Read a two-page summary of the findings here

Nicholls, E., Griffiths-Lee, J., Basu, P., Chatterjee, S. & Goulson, D. (2023) Crop-pollinator interactions in urban and peri-urban farms in the UK. Plants, People, Planet.

Griffiths-Lee, J., Nicholls, E., & Goulson, D. (2023) Sow Wild! Effective methods and identification bias in pollinator-focused experimental citizen science. Citizen Science, Theory and Practice.

Nicholls, E., Rands, S., Botias, C., Hempel de Ibarra, N. (2022) Flower sharing and pollinator health: a behaviour perspective. Philosophical Transactions of the Royal Society B, 377, 20210157.

Griffiths-Lee, J. Davenport, B., Foster, B., Nicholls, E., & Goulson, D. (2022) Sown wildflowers between vines increase beneficial insect abundance and richness in a British vineyard. Agricultural and Forest Entomology

Griffiths-Lee, J., Goulson, D. & Nicholls, E. (2022) Grape expectations: a survey of British vineyard land management practices from an environmental perspective. Journal of Agriculture and Sustainability, 15. 

Griffiths-Lee, J., Nicholls, E., & Goulson, D. (2022). Sown mini-meadows increase pollinator diversity in gardens. Journal of Insect Conservation, 1-16.

Goulson, D., & Nicholls, E. (2022). Anthropogenic influences on bee foraging. Science375(6584), 970-972.

Nicholls, E., Rossi, M., & Niven, J. E. (2021). Larval nutrition impacts survival to adulthood, body size, and the allometric scaling of metabolic rate in adult honeybees. Journal of Experimental Biology, 224, jeb242393.

Nicholls, E., Ely, A., Birkin, L., Basu, P., & Goulson, D. (2020). The contribution of small-scale food production in urban areas to the sustainable development goals: A review and case study. Sustainability Science, 1-15.

Griffiths‐Lee, J., Nicholls, E., & Goulson, D. (2020). Companion planting to attract pollinators increases the yield and quality of strawberry fruit in gardens and allotments. Ecological Entomology45(5), 1025-1034.

Nicholls, E., Krishna, S., Wright, O., Stabler, D., Krefft, A., Somanathan, H., & de Ibarra, N. H. (2019). A matter of taste: the adverse effect of pollen compounds on the pre-ingestive gustatory experience of sugar solutions for honeybees. Journal of Comparative Physiology A205(3), 333-346.

Nicholls, E., Botias, C., Rotheray, E. L., Whitehorn, P., David, A., Fowler, R., ... & Goulson, D. (2018). Monitoring neonicotinoid exposure for bees in rural and peri-urban areas of the UK during the transition from pre-to post-moratorium. Environmental Science & Technology, 52(16), 9391-9402.

Balamurali, G. S., Nicholls, E., Somanathan, H., & de Ibarra, N. H. (2018). A comparative analysis of colour preferences in temperate and tropical social bees. The Science of Nature105(1-2), 8.

Nicholls, E., Fowler, R., Niven, J. E., Gilbert, J. D., & Goulson, D. (2017). Larval exposure to field-realistic concentrations of clothianidin has no effect on development rate, over-winter survival or adult metabolic rate in a solitary bee, Osmia bicornisPeerJ5, e3417.

Nicholls, E., & Hempel de Ibarra, N. (2017). Assessment of pollen rewards by foraging bees. Functional Ecology, 31(1), 76-87.

Goulson, D., Nicholls, E., Botías, C., & Rotheray, E. L. (2015). Bee declines driven by combined stress from parasites, pesticides, and lack of flowers. Science347(6229).

Nicholls, E. K., Ehrendreich, D., & Hempel de Ibarra, N. (2015). Differences in color learning between pollen-and sucrose-rewarded bees. Communicative & Integrative Biology8(4), e1052921.

Nicholls, E., & de Ibarra, N. H. (2014). Bees associate colour cues with differences in pollen rewards. Journal of Experimental Biology217(15), 2783-2788.

Nicholls, E., & De Ibarra, N. H. (2013). Pollen elicits proboscis extension but does not reinforce PER learning in honeybees. Insects4(4), 542-557.